North American Lepidoptera Biodiversity LLC


Contributions to a Check List of the Lepidoptera of Fallen Timbers Environmental Center in Outagamie County, Wisconsin

(Macrolepidoptera Families: Thyatiridae, Drepanidae, Geometridae, Epiplemidae, Hesperiidae, Papilionidae, Pieridae, Lycaenidae, Riodinidae, Nymphalidae, Mimallonidae, Lasiocampidae, Apatelodidae, Saturniidae, Sphingidae, Notodontidae, and Noctuidae; also including: Hepialidae, Cossidae, Attevidae, Yponomeutidae, Zygaenidae, Limacodidae, and Thyrididae)


By Hugo L. Kons Jr.


Posted on the Web 8 September 2007


Most Recent Update: Feb. 2014


Note: While this check list does not contain detailed information for the included species, an upcoming volume of the North American Journal of Lepidoptera Biodiversity will present phenology data, habitat data and analyses, hypotheses of habitat dependency for many of the included species, and other information and analyses. To receive an e-mail notification when this volume becomes available, contact the author at


Abstract: I present a check list of 653 Lepidoptera species recorded from Fallen Timbers Environmental Center in Outagamie County, Wisconsin, including 642 species of Macrolepidoptera.

Introduction: Fallen Timbers Environmental Center (N 4428.7' W 8824.6') contains an extensive inland wetland hardwood forest with many open to shrubby wetland and sedge meadow openings. Also present are sizable mesic meadows and fields as well as a man-made pond. I have conducted extensive diurnal and nocturnal field work at this locality.

I conducted surveys at Fallen Timbers primarily from 1989-1996, but I also have a small number of records from earlier. Surveys were conducted from 1989-1991 with bait traps and diurnal searching. From 1992-1995 (and once in 1996) I surveyed Fallen Timbers at night with UV lights and bait trails, in addition to continuing to conduct diurnal surveys.

I first produced check lists of Fallen Timbers Rhopalocera and nocturnal Macrolepidoptera in the early 1990s, but only as unpublished reports. These check lists have been continuously updated over the years and the current check list probably approaches a thorough inventory of the species I have documented in the included families, although I still have backlog material. In addition, some of the detailed survey data for Fallen Timbers appears in a variety of unpublished reports which have been distributed to land managers, the WI Department of Natural Resources, and various Lepidopterists. This internet site is my first paper on Fallen Timbers Lepidoptera that is more widely available, but more detailed papers are planned for future volumes of the North American Journal of Lepidoptera Biodiversity.


Methods: My surveys were conducted with UV sheets, UV traps, type P and F bait traps, bait trails, and diurnal collecting with nets and jars. Follow this link for illustrations and a discussion of these survey methods.


Voucher Specimens: At least one voucher specimen of each species I report from Fallen Timbers is currently housed in my personal research collection, with the exception of Catocala subnata (housed in the research collection of Robert J. Borth). In addition, I have made donations of Fallen Timbers Lepidoptera to the Milwaukee Public Museum, and Robert Borth has collected some material from Fallen Timbers as well. I identified or verified specimens of all of the species I list from Fallen Timbers with the exception of some Eupithecia (determined by George Balogh).


Classification: The classification primarily follows the Hodges et al. (1983) check list with the exception of Nymphalidae (concept of Harvey 1991) and Noctuidae, incorporating information from Poole (1995), Mitchell et al. (2006), and Zahiri et al. (2011, 2012a, 2012b, 2013). For a more detailed discussion of adopted changes in classification from the Hodges et al. (1983) check list, see the Classification section of the Kons, Borth, and Barina Check List of Wisconsin Macrolepidoptera. For those species listed in Hodges et al. (1983), I include the original check list number, which serves as a citation for the describing author and year of description.


Results: To date I can confirm the documented occurrence of 653 Lepidoptera species in the included families from Fallen Timbers Environmental Center, including 642 species of Macrolepidoptera. These species are listed in the check list at the end of this report. All but a few of the species on the following list were collected during my surveys of Fallen Timbers from 1989-1995. The exceptions include Gaeides xanthioides (1983), Oligia hausta (1996), and Catocala subnata (collected in 2007 by Robert J. Borth).


Discussion: Fallen Timbers Environmental Center has one of the longest lists of documented Macrolepidoptera species among the eastern North America localities of comparable size for which I have compiled check lists or seen check lists in other publications. However, usually the most important factor influencing relative differences in the lengths of documented Lepidoptera species lists between different sites is the amount of survey effort, rather than differences in species diversity in nature. The only sites that I have surveyed as or more extensively than Fallen Timbers (with surveys conducted over multiple seasons and covering the full flight season when adult Lepidoptera are active) are Summer Street in Appleton (Outagamie County, Wisconsin), Mosquito Hill Nature Center (Outagamie County, Wisconsin), the American Entomological Institute property (Alachua County, Florida) and Southwest 20th Avenue in Gainesville (Alachua County, Florida). Among these localities, Fallen Timbers has the second highest species list (nearby Mosquito Hill Nature Center has the longest). Another locality in the Kons-Borth-Barina data set with a longer Macrolepidoptera species list is the barrens and oak-pine forests northwest of Dunbar. This site was studied less extensively than Fallen Timbers, and thus probably really does have a more diverse Macrolepidoptera fauna. The Fallen Timbers Macrolepidoptera species list contains about 46.3% of the verified Macrolepidoptera species listed in the Kons, Borth, and Barina Check List of Wisconsin Macrolepidoptera, or 44.1% of WI Macrolepidoptera species if I include literature records which I have not verified.

Fallen Timbers is a very significant locality in terms of hosting populations of wetland dependent Lepidoptera species. An upcoming NAJLB Volume on Wisconsin Lepidoptera by Kons, Borth, and Barina, includes analyses and preliminary lists of Wisconsin Macrolepidoptera species hypothesized to be dependent on particular habitat types, including wetlands and particular types of wetlands such as hydric hardwood forest and sedge meadow. Potentially wetland dependent species recorded from Fallen Timbers include: Sthenopis argenteomaculatus, Gueneria similaria (hydric hardwood forest), Scopula cacuminaria, Scopula quadrilineata, Plemyria georgeii, Heterophleps triguttaria (hydric hardwood forest), Carterocephalus palemon, Ancyloxpha numitor, Poanes massasoit (sedge meadow), Poanes viator (sedge meadow), Euphyes dion (sedge meadow), Euphyes conspicua (sedge meadow), Hyllolycaena hyllus, Satyrium acadicum, Clossiana selene (primarily sedge meadow), Chlosyne harrissii (sedge meadow), Satyrodes eurydice (sedge meadow), Satyrodes appalachia (hydric hardwood forest), Macrochilo absorptalis, Macrochilo bivittata, Phalaenostola hanhami, Hypenodes caducus, Dyspyralis nigella (hydric hardwood forest), Ledaea perditalis, Catocala crataegi (hydric hardwood forest), Catocala alabamae (not range wide), Nola cilicoides, Capis curvata, Plusia putnami (sedge meadow), Plusia contexta (sedge meadow), Plusia venusta (sedge meadow), Acronicta oblinata, Apamea nigrior (hydric hardwood forest), Luperina stipata, Eremobina jocasta (sedge meadow), Meropleon diversicolor, Lemmeria digitalis (sedge meadow), Chortodes inquinata (sedge meadow), Chortodes defecta (sedge meadow), Archanara oblonga (sedge meadow), Papaipema cataphracta (sedge meadow), Papaipema impecuniosa (sedge meadow), Papaipema nepheleptena (sedge meadow), Papaipema eupatorii (sedge meadow), Papaipema unimoda, Bellura obliqua, Lithophane oriunda, Xanthia togata, Catabena lineolata, Leucania linita (sedge meadow), and Agrotis gladiaria (sedge meadow).

It is also noteworthy that some wetland dependent Lepidoptera may be less susceptible to catastrophic flooding events at a locality like Fallen Timbers than populations occurring in riparian floodplains. Poanes viator was common in Mosquito Hill Nature Center sedge meadows on the Wolf River floodplain from 1989-1992, and Euphyes dion was present there also but uncommon. This area is normally flooded in the spring, but in 1993 (an usually wet year with major flooding problems along the Mississippi) the area remained flooded into July and August and the sedge meadow community never grew. The following year the sedge meadow vegetation returned, but surveys in 1994 and 1995 did not uncover any P. viator or E. dion. In contrast, while the inland swamp forest/sedge meadow complexes at Fallen Timbers had unusually high water levels in 1993, the sedge meadow plant communities and associated butterflies (Poanes viator, Poanes massasoit, Euphyes dion, Euphyes conspicua, Chlosyne harrisii, Clossiana selene, and Satyrodes eurydice) all survived.



Acknowledgments: I thank Libby Dorn, director of Fallen Timbers Environmental Center, for her cooperation and encouragement with my Lepidoptera surveys at the environmental center. The authorization she provided to collect specimens with 24 hour access to the environmental center was crucial to the success of this project. I am especially grateful to my parents, Hugo & Sharon Kons Sr., for much support and tolerance for my research activities over many years. I thank my colleagues Robert Borth and Thomas Barina for their collaboration in our study of Wisconsin Lepidoptera. I give special thanks to Ken Hall of the University of Wisconsin-Stevens Point (retired) for his support and encouragement of my undergraduate Lepidoptera research activities. I thank the University of Wisconsin-Stevens Point and the American Entomological Institute for lab space and infrastructural support. George Balogh examined some of the first Geometrid specimens I collected at Fallen Timbers and was instrumental in helping me start my research collection for this family. George Balogh also determined specimens for all of the Fallen Timbers Eupithecia species. Hugo and Sharon Kons Sr. assisted with building light and bait traps, and Richard Merkhofer provided the initial design for the inverted cone platform bait trap. For facilitating access to museum collections I thank Donald Harvey, Patricia Poole, and Robert Robbins (National Museum of Natural History) and Su Borkin (Milwaukee Public Museum). Also, for assistance with acquiring chemicals important to collecting or genitalic dissections, I thank Niklaus Hostettler, Jim Lloyd, Robert Robbins, and David Wahl.


Check List of the Lepidoptera of Fallen Timbers Environmental Center


Species Total for Included Families 653







6235 Habrosyne scripta

6236 Habrosyne gloriosa

6237 Pseudothyatira cymatophoroides



Drepaninae 3

6251 Drepana arcuata

6252 Drepana bilineata

6253 Eudeilinia herminiata

Oretinae 1

6255 Oreta rosea






Archierinae 1

6256 Archieris infans


Ennominae 83

6261 Heliomata cycladata

6270 Proitame virginalis

6273 Itame pustularia

6278 Itame evagaria

6290 Itame loricaria

6303 Itame subcessaria

6326 Semiothisa aemulataria

6330 Semiothisa ulsterata

6342 Semiothisa bisignata

6351 Semiothisa oweni

6362 Semiothisa continuata

6386 Semiothisa ocellinata

6397 Semiothisa mellistrigata

6405 Semiothisa gnophosaria

6436 Ematurga amitaria

6570 Aethalura intertexta

6583 Iridopsis ephyraria

6584 Iridopsis humaria

6588 Iridopsis larvaria

6590 Anavitrinelia pampinaria

6597 Ectropis crepuscularia

6598 Protoboarmia porcelaria

6620 Melanolophia canadaria

6621 Melanolophia signataria

6637 Eufidonia convergaria

6640 Biston betularia

6651 Lycia ursaria

6654 Hypagyrtis unipunctata

6658 Phigalia titea

6660 Phigalia strigitaria

6662 Paleacrita vernata

6665 Erannis tiliaria

6666 Lomographa semiclarata

6667 Lomographa vestaliata

6668 Lomographa glomeraria

6677 Cabera erythemaria

6678 Cabera variolaria

6720 Lytrosis unitaria

6724 Euchlaena serrata

6726 Euchlaena obtusaria

6728 Euchlaena effecta

6729 Euchlaena johnsonaria

6734 Euchlaena marginaria

6740 Xanthotype urticaria

6743 Xanthotype sospeta

6753 Pero honestaria

6754 Pero hubneraria

6755 Pero morrisonaria

6763 Nacophora quernaria

6796 Campaea perlata

6797 Ennomos magnaria

6798 Ennomos subsignaria

6799 Epirranthis substriataria

6804 Petrophora subaequaria

6807 Tacparia detersata

6811 Homochlodes lactispargaria

6812 Homochlodes fritillaria

6815 Gueneria similaria

6819 Metanema inatomaria

6820 Metanema determiniata

6822 Metarranthis duaria

6825 Metarranthis indeclinata

6827 Metarranthis refractaria

6835 Cepphis armataria

6836 Anagoga pulveraria occiduaria

6837 Probole alienaria

6838 Probole amicaria

6839.1 Probole nyssaria

6842 Plagodis phlogosaria

6843 Plagodis fervidaria

6844 Plagodis alcoolaria

6884 Besma endropiaria

6885 Besma quercivoraria

6888 Lambdina fiscellaria

6906 Nepytia canosaria

6912 Sicya macularia

6941 Eusarca confusaria

6963 Tetracis crocallata

6964 Tetracis cachexiata

6965 Eugonobapta nivosaria

6966 Eutrapela clemataria

6982 Prochoerodes lineola

7009 Nematocampa resistaria


Geometrinae 3

7048 Nemoria mimosaria

7058 Synchlora aerata

7071 Chlorochlamys chloroleucaria


Sterrhinae 7

7132 Pleuroprucha insulsaria

7139 Cyclophora pendulinaria

7146 Haematopis grataria

7157 Scopula cacuminaria

7159 Scopula limboundata

7165 Scopula quadrilineata

7169 Scopula inductata


Larentiinae 40

7196 Eulithis diversilineata

7197 Eulithis gracilineata

7201 Eulithis testata

7205 Eulithis flavibrunneata

7206 Eulithis explanata

7216 Plemyria georgeii

7236 Hydriomena renunciata

7237 Hydriomena transfigurata

7254 Hydriomena ruberata

7285 Triphosa haesitata

7290 Coryphista meadii

7292 Hydria prunivorata

7293 Rheumaptera hastata

7307 Mesoleuca ruficillata

7312 Spargania magnoliata

7329 Anticlea vasiliata

7230 Anticlea multiferata

7368 Xanthorhoe labradorensis

7388 Xanthorhoe ferrugata

7390 Xanthorhoe lacustrata

7394 Epirrhoe alternata

7399 Euphyia intermediata

7414 Orthonama obstipata

7416 Orthonama centrostrigaria

7419 Hydrelia lucata

7423 Hydrelia albifera

7428 Venusia comptaria

7430 Trichodezia albovittata

7433 Epirrita autumnata

7440 Eubaphe mendica

7445 Horisme intestinata

7474 Eupithecia miserulata

7491 Eupithecia fletcherata

7518 Eupithecia intricata

7529 Eupithecia absinthiata

7605 Eupithecia ravocostaliata

7640 Lobophora nivigerata

7645 Heterophleps refusaria

7647 Heterophleps triguttaria

7648 Dyspteris abortivaria



7653 Calledapteryx dryopterata





Pyrginae 4

3945 Erynnis icelus

3947 Erynnis juvenalis

3966 Pyrgus communis

3977 Pholisora catullus


Heteropterinae 1

3982 Carterocephalus palemon


Hesperiinae 19

4004 Ancyloxpha numitor

4012 Thymelicus lineola

4033 Hesperia sassacus

4036 Polites coras

4041 Polites themistocles

4043 Polites mystic

4047 Wallengrenia egeremet

4048 Pompeius verna

4049 Atalopedes campestris

4051 Atrytone logan

4058 Poanes massasoit

4059 Poanes hobomok

4064 Poanes viator

4072 Euphyes dion

4075 Euphyes conspicua

4078 Euphyes vestris

4096 Amblyscirtes hegon

4105 Amblyscirtes vialis

4111 Lerodea eufala





4159 Papilio polyxenus

4176 Papilio glaucus

4176.1 Papilio canadensis



4195 Pieris napi

4197 Pieris rapae

4209 Colias philodice

4210 Colias eurytheme

4224 Zerene cessonia



4249 Feniseca tarquinius

4253 Gaeides xanthoides

4256 Hyllolycaena hyllus

4285 Satyrium liparops

4282 Satyrium calanus

4281 Satyrium edwardsii

4278 Satyrium acadicum

4326 Callophrys henrici

4361 Everes comyntas

4363 Celastrina ladon [complex]



4420 Polygonia interrogationis

4421 Polygonia comma

4429 Polygonia progne

4430 Nymphalis vau-album

4432 Nymphalis antiopa

4433 Aglais milberti

4434 Vanessa virginiensis

4435 Vanessa cardui

4437 Vanessa atlanta

4440 Junonia coenia

4450 Speyeria cybele

4451 Speyeria aphrodite

4459 Speyeria atlantis

4464 Clossiana selene

4465 Clossiana bellona

4481 Phyciodes tharos

4481.1 Phyciodes pascoensis

4482 Phyciodes batesii

4489 Chlosyne gorgone

4491 Chlosyne harrisii

4522 Basilarchia arthemis

4523 Basilarchia archippus

4557 Astereocampa celtis

4568.1 Enodia anthedon

4568.3 Satyrodes eurydice

4569 Satyrodes appalachia

4578 Megisto cymela

4587 Cercyonis pegala

4614 Danaus plexippus





Macromphaliinae 2

7670 Tolype velleda

7673 Tolype laricis

Gastropachinae 1

7687 Phyllodesma americana

Lasiocampinae 2

7698 Malacosoma disstria

7701 Malacosoma americanum





7665 Olceclostera angelica



Citheroniinae 1

7715 Dryocampa rubicunda

Saturniinae 3

7757 Antheraea polyphemus

7758 Actias luna

7767 Hyalophora cecropia




Sphinginae 12

7784 Dolba hyloeus

7787 Ceratomia undulosa

7796 Sphinx eremitus

7807 Sphinx canadensis

7809 Sphinx kalmiae

7810 Sphinx gordius (incl. poecile)

7811 Sphinx luscitiosa

7821 Smerinthus jamaicensis

7822 Smerinthus cerisyi

7824 Paonias excaecatus

7825 Paonias myops

7828 Pachysphinx modesta


Macroglossinae 8

7853 Hemaris thysbe

7855 Hemaris diffinis

7870 Sphecodina abbottii

7871 Deidamia inscripta

7873 Amphion floridensis

7885 Darapsa myron

7886 Darapsa pholus

7894 Hyles lineata





7895 Clostera albosigma

7898 Clostera strigosa

7901 Clostera apicalis

7902 Datana ministra

7906 Datana contractata

7907 Datana integerrima

7908 Datana perspicua

7915 Nadata gibbosa

7917 Hyperaeschra georgica

7920 Peridea angulosa

7922 Pheosia rimosa

7924 Odontosia elegans

7928 Notodonta simplaria

7929 Nerice bidentata

7930 Ellida caniplaga

7931 Gluphisia septentrionis

7933 Gluphisia avimacula

7939 Furcula occidentalis

7952 Symmerista canicosta

7953 Symmerista leucitys

7975 Macrurocampa marthesia

7983 Heterocampa obliqua

7994 Heterocampa guttivitta

7999 Lochmaeus bilineata

8005 Schizura ipomoeae

8006 Schizura badia

8007 Schizura unicornis

8011 Schizura leptinoides

8012 Oligocentria semirufescens




Herminiinae 32

8322 Idia americalis

8323 Idia aemula

8323.1 Idia concisa

8324 Idia majoralis

8326 Idia rotundalis

8328 Idia julia

8334 Idia lubricalis

8338 Phalaenophana pyramusalis

8340 Zanclognatha lituralis

8345 Zanclognatha laevigata

8348 Zanclognatha pedipilalis cpx. A

8349 Zanclognatha protumnusalis

8351 Zanclognatha cruralis cpx. sp A

8352 Zanclognatha marcidilinea (formerly jacchusalis)

8353 Zanclognatha ochreipennis

8355 Chytolita morbidalis

8356 Chytolita petrealis

8357 Macrochilo absorptalis

8357.1 Macrochilo hypocritialis Fgn.

8358 Macrochilo litophora

8359 Macrochilo bivittata

8360 Macrochilo orciferalis

8361 Macrochilo louisiana

8362 Phalaenostola metonalis

8363 Phalaenostola eumelusalis

8364 Phalaenostola larentioides

8365 Phalaenostola hanhami

8370 Bleptina caradrinalis

8379 Renia factiosalis

8384.1 Renia flavipunctalis

8393 Lascoria ambigualis

8397 Palthis angulalis


Rivulinae 1

8404 Rivula propinqualis


Hypenodinae 2

8420 Hypenodes caducus

Hypenodes sp. nr. fractilinea


Subfamily Unknown 1

8428 Dyspyralis nigella


Hypeninae 11

8441 Hypena manalis

8442 Hypena baltimoralis

8443 Hypena bijugalis

8445 Hypena abalienalis

8446 Hypena deceptalis

8448 Hypena sordidula

8450 Hypena atomaria

8452 Hypena edictalis

8455 Hypena eductalis

8461 Hypena humuli

8465 Hypena scabra


Subfamily Unkown 1

8479 Spargaloma sexpunctata


Pangraptinae 1

8491 Ledaea perditalis


Boletobiinae 1

8500 Metalectra quadrisignata


Calpinae 1

8536 Calyptra canadensis


Scoliopteryginae 1

8555 Scoliopteryx libatrix


Eulepidotinae 1

8574 Anticarsia gemmatalis


Erebinae (=Catocalinae) 31

8592 Cissusa spadix

8689 Zale lunata

8692 Zale galbanata

8695 Zale undularis

8697 Zale minerea

8713.1 Zale intenta (formerly lunifera)

8717 Zale horrida

8719 Euparthenos nubilis

8727 Parallelia bistriaris

8738 Caenurgina crassiuscula

8739 Caenurgina erechtea

8788 Catocala retecta

8797 Catocala subnata [Col. Robert J. Borth]

8798 Catocala neogama

8801 Catocala ilia

8802 Catocala cerogama

8803 Catocala relicta

8805 Catocala unijuga

8806 Catocala parta

8817 Catocala briseis

8822 Catocala meskei

8832 Catocala cara

8833 Catocala concumbens

8834 Catocala amatrix

8851 Catocala coccinata

8857 Catocala ultronia

8858 Catocala crataegi

8863 Catocala mira

8864 Catocala grynea

8867 Catocala blandula

8869 Catocala alabamae


Arctiinae 25

8043 Eilema bicolor

8045.1 Crambidia pallida

8090 Hypoprepia fucosa

8098 Clemensia albata

8109 Haploa reversa

8111 Haploa lecontei

8112 Haploa confusa

8114 Holomelina laeta

8121 Holomelina aurantiaca

8123 Holomelina ferruginosa

8129 Pyrrharctia isabella

8131 Estigmene acrea

8134 Spilosoma congrua

8137 Spilosoma virginica

8140 Hyphantria cunea

8156 Phragmatobia fuliginosa

8175 Grammia virguncula

8196 Grammia parthenice

8197 Grammia virgo

8203 Halysidota tessellaris

8230 Cycnia tenera

8231 Cycnia oregonensis

8238 Euchaetes egle

8262 Ctenucha virginica

8267 Cisseps fulvicollis

Lymantriinae 4

8300 Dasychira cinnamomea

8302 Dasychira obliquata

8314 Orgyia definita

8316 Orgyia leucostigma


Euteliinae 1

8955 Marathyssa inficita


Risobinae 3

8969 Baileya doubledayi

8970 Baileya ophthalmica

8971 Baileya dormitans


Chloephorinae 1

8975 Nycteola frigidana

Nolinae 4

8983 Meganola miniscula

8983.2 Meganola spodia

8990 Nola cilicoides

8992 Nola triquetrana


Subfamily Unknown 1

9038 Hyperstrotia villificans


Eustrotiinae 8

9046 Lithacodia bellicula

9047 Lithacodia muscosula

9048 Lithacodia albidula

9051 Lithacodia musta

9049 Maliattha synochitis

9053 Pseudostrotia carneola

9284 Anterastria teratophora

9059 Capis curvata

Condicinae 7

9690 Condica videns

9696 Condica vecors

9720 Ogdoconta cinereola

9057 Homophoberia apicosa

9065 Leuconycta diphteroides

9066 Leuconycta lepidula

9661 Crambodes talidiformis


Plusiinae 15

8881 Abrostola urentis

8887 Trichoplusia ni

8890 Pseudoplusia includens

8896 Diachrysia aeroides

8897 Diachrysia balluca

8898 Allagrapha aerea

8905 Eosphoropteryx thyatyroides

8907 Megalographa biloba

8908 Autographa precationis

8924 Anagrapha falcifera

8927 Syngrapha epigaea

8950 Plusia putnami

Plusia magnimacula

8952 Plusia contexta

8953 Plusia venusta


Acontiinae 2

9090 Tarachidia candefacta

9095 Tarachidia erastrioides

Amphipyrinae 3

9638 Amphipyra pyramidoides

9639 Amphipyra tragopoginis

9640 Amphipyra glabella


Eriopinae 1

9631 Callopistria mollissima

Psaphidinae 2

10019 Psaphida resumens

10021 Copivaleria grotei


Heliothinae 4

11068 Helicoverpa zea

11128 Schinia arcigera

11135 Schinia rivulosa

11149 Schinia trifascia

Agaristinae 2

9299 Eudryas unio

9301 Eudryas grata


Pantheinae 2

9185 Colocasia propinquilinea

9189 Charadra deridens


Raphiinae 1

9193 Raphia frater


Acronictinae 26

9062 Cerma cerintha

9200 Acronicta americana

9203 Acronicta dactylina

9205 Acronicta lepusculina

9206 Acronicta leporina

9207 Acronicta innotata

9212 Acronicta grisea

9225 Acronicta vinnula

9226 Acronicta superans

9229 Acronicta hasta

9236 Acronicta morula

9237 Acronicta interrupta

9238 Acronicta lobeliae

9241 Acronicta fragilis

9243 Acronicta ovata

9244 Acronicta modica

9245 Acronicta haesitata

9250 Acronicta inclara

9251 Acronicta retardata

9257 Acronicta impleta

9259 Acronicta noctivaga

9261 Acronicta impressa

9272 Acronicta oblinata

9280 Simyra henrici sp. B

9281 Agriopodes fallax

9286 Harrisimemna trisignata

Cucullinae 3

10197 Cucullia florea

10200 Cucullia asteroides

10202 Cucullia convexipennis


Noctuinae 171

9328 Apamea nigrior

9333 Apamea lignicolora

9348 Apamea amputatrix

9351 Apamea alia

9364 Apamea sordens

9367 Apamea dubitans

9373 Apamea helva

9382 Apamea devastator

9393 Luperina stipata

9398 Eremobina jocasta

9404 Oligia modica

9408 Oligia exhausta

9409 Oligia hausta/semicana cmplx.

9413 Oligia tonsa

9419 Oligia mactata

9427 Meropleon diversicolor

9429 Lemmeria digitalis

9431 Parastichtis suspecta

9433 Xylomoia chagnoni

9437 Chortodes inquinata

9443 Chortodes defecta

9449 Archanara oblonga

9453 Celaena reniformis

9454 Amphipoea velata

9456 Amphipoea interoceanica

9457 Amphipoea americana

9466 Papaipema cataphracta

Papaipema sp.

9471 Papaipema arctivorens

9473 Papaipema impecuniosa

9480 Papaipema pterisii

9483 Papaipema inquaesita

9486 Papaipema birdi

9490 Papaipema nepheleptena

9495 Papaipema furcata

9497 Papaipema necopina

9501 Papaipema eupatori

9509 Papaipema unimoda

9514 Hydraecia micacea

9525 Bellura obliqua

9545 Euplexia benesimilis

9546 Phlogophora iris

9547 Phlogophora periculosa

9549 Enargia decolor

9550 Enargia infumata

9551 Enargia mephisto

9555 Ipimorpha pleonectusa

9556 Chytonix palliatricula

9560 Dypterygia rozmani

9578 Hyppa xylinoides

9582 Nedra ramosula

9626 Trachea delicata

9647 Athetis miranda

9649 Athetis mendosa

9662 Balsa malana

9663 Balsa tristrigella

9664 Balsa labecula

9666 Spodoptera frugiperda

9669 Spodoptera ornithogalli

9671 Spodoptera dolichos

9678 Elaphria versicolor

9681 Elaphria festivoides

9688 Galgula partita

9815 Cosmia calami

9818 Amolita fessa

9874 Xylena curvimacula

9878 Lithomoia germana

9886 Lithophane patefacta

9887 Lithophane bethunei

9891 Lithophane amanda

9892 Lithophane disposita

9893 Lithophane hemina

9894 Lithophane oriunda

9910 Lithophane antennata

9914 Lithophane laticinerea

9915 Lithophane grotei

9916 Lithophane unimoda

9917 Lithophane fagina

9922 Lithophane pexata

9929 Pyreferra hesperidago

9930 Pyreferra citromba

9933 Eupsilia vinulenta

9935 Eupsilia tristigmata

9936 Eupsilia morrisoni

9939 Eupsilia devia

9942 Metaxaglaea inulta

9950 Chaetaglaea sericea

9952 Eucirroedia pampina

9957 Sunira bicolorago

9961 Anathix ralla

9962 Anathix puta

9965 Xanthia togata

9998 Brachylomia algens

10033 Catabena lineolata

10223 Discestra trifolii

10272 Mamestra curialis

10276 Polia imbrifera

10280 Polia purpurissata

10288 Polia detracta

10292 Melanchra adjuncta

10293 Melanchra picta

10295 Melanchra assimilis

10297 Lacanobia atlantica

10299 Lacanobia subjuncta

10300 Spirameter grandis

10301 Spirameter lutra

10304 Trichordestra legitima

10368 Lacinipolia meditata

10397 Lacinipolia renigera

10405 Lacinipolia lorea

10406 Lacinipolia olivacea

10431 Faronta diffusa

10436 Aletia oxygala

10438 Pseudaletia unipuncta

10440 Leucania linita

10444 Leucania phragmatidicola

10445 Leucania linda

10446 Leucania multilinea

10446.1 Leucania lapidaria

10447 Leucania commoides

10462 Leucania pseudargyria

10487 Orthosia rubescens

10491 Orthosia alurina

10495 Orthosia hibisci

10501 Crocigrapha normani

10513 Egira dolosa

10518 Achatia distincta

10520 Morrisonia evicta

10291 Morrisonia latex

10524 Nephelodes minians

10532 Homorthodes furfurata

10563 Protorthodes oviduca

10578 Pseudorthodes vecors

10585 Orthodes crenulata

10289 Orthodes goodelli

10597 Orthodes cynica

10648 Agrotis gladiaria

10651 Agrotis venerabilis

10663 Agrotis ipsilon

10670 Feltia jaculifera

10674 Feltia subgothica

10675 Feltia tricosa

10676 Feltia herilis

10705 Euxoa messoria

10793 Euxoa scholastica

10803 Euxoa velleripennis

10805 Euxoa tessellata

10903 Euagrotis illapsa

10915 Peridroma saucia

10891 Ochropleura implecta

10926 Spaelotis clandestina

10928 Graphiphora augur

10929 Eurois occulta

10942 Xestia c-nigrum

10942.1 Xestia dolosa

10943 Xestia normaniana

10944 Xestia smithii

10988 Coenophila opacifrons

10955 Agnorisma badinodis

10950 Pseudohermonassa bicarnea

10951 Pseudohermonassa tenuicula

10994 Cerastis tenebrifera

10996 Cerastis salicarum

11000 Anaplectoides prasina

11001 Anaplectoides pressus

11006 Protolampra brunneicollis

11007 Eueretagrotis sigmoides

11008 Eueretagrotis perattenta

11012 Cryptocala acadiensis

11029 Abagrotis alternata

11043 Abagrotis cupida


Some Other Lepidoptera Families



18 Sthenopis argenteomaculatus


2676 Acossus populi


2401 Atteva punctella



4624 Harrisina americana


4652 Tortricidia testacea

4659 Packardia geminata

4665 Lithacodes fasciola

4667 Apoda y-inversum

4668 Apoda biguttata

4697 Euclea delphini


6076 Thyris maculata


References Cited


Harvey, Donald J. 1991. Higher Classification of the Nymphalidae. Appendix B in The Development and Evolution of Butterfly Wing Patterns by H. F. Nijhout. Smithsonian Institution Press.

Hodges, Ronald W. et. al. 1983. Check List of the Lepidoptera of America North of Mexico. Great Britian, University Press, Cambridge. 284 pp.

Mitchell, Andrew, Charles Mitter, and Jerome C. Regier. 2006. Systematics and evolution of the cutworm moths (Lepidoptera: Noctuidae): evidence from two protein-coding nuclear genes. Systematic Entomology, 31: 21-46.

Poole, Robert W. 1995. Noctuoidea, Noctuidae (part) in Dominick, R. B. et al., The Moths of America North of Mexico, Fasc. 26.1.

Zahiri, Reza, Ian J. Kitching, J. Donald Lafontaine, Marko Mutanen, Lauri Kaila, Jeremy Holloway & Niklas Wahlberg. 2011. A new molecular phylogeny offers hope for a stable family level classification of the Noctuouidea (Lepidoptera). Zoologica Scripta.

Zahiri, Reza, Jeremy Holloway, Ian J. Kitching, J. Donald Lafontaine, Marko Mutanen, & Niklas Wahlberg. 2012a. Molecular phylogenetics of Erebidae (Lepidoptera: Noctuoidea). Systematic Entomology, 37: 102-124.

Zahiri, Reza, J. Donald Lafontaine, Jeremy Holloway, Ian J. Kitching, B. Christian Schmidt, Lauri Kaila, & Niklas Wahlberg. 2012b. Major lineages of Nolidae (Lepidoptera, Noctuoidea) elucidated by molecular phylogenetics. Cladistics 1 (2012): 1-23.

Zahiri, Reza, J. Donald Lafontaine, Christian Schmidt, Jeremy Holloway, Ian J. Kitching, Marko Mutanen, & Niklas Wahlberg. 2013. Relationships among the basal lineages of Noctuidae (Lepidoptera, Noctuoidea) based on eight gene regions. Zoologica Scripta.